John G. Costouros, MD

Professional Education

• Internship: University of California San Francisco CA (1999)
• Medical Education: University of California San Francisco CA (1998)
• Fellowship: Harvard Medical School MA (2005)
• Fellowship, University of Zurich, Balgrist Hospital Shoulder and Elbow Surgery (2005)
• Residency: University Of California, San Francisco CA (2004)
• Fellowship, University of California, San Francisco (UCSF), Department of Orthopaedic Surgery Cartilage Injury, Apoptosis (2002)
• B.S. with Honors, Stanford University Biological Sciences (1994)